Reprints of publications are intended for personal use only.


Fonseca EM, Pelletier TA, Decker SK, Parsons DJ, Carstens BC. Pleistocene glaciations contributed to the latitudinal gradient of within-species genetic diversity. Evolution Letters, in press.

Smith ML, C Lang, D Sneddon, J Wallace, AM Rankin, A Espíndola, DC Tank, J Sullivan, BC Carstens. Comparative Phylogeography of Leaf-litter dwelling invertebrates from the Pacific Northwest. Journal of Northwest Science 96(2), in press.

Wieringa JG, Nagel J, Campbell CJ, Nelson DM, Carstens BC, Gibbs HL. Combining stable isotope, trace element, and species distribution models to assess the geographic origins of migratory bats. Ecosphere 14, e4588.

Rao A, Baken L, Lubertozzi A, Jahan I, Tedeschi F, Jankowsky E, Carstens BC, Poss KD, Basking KK, Goldman JA. The Translation Initiation Factor Homolog, eif4e1c, Regulates Cardiomyocyte Metabolism and Proliferation During Heart Regeneration. Development 150: dev 2011376.

Pérez-Alquicira J, Wehncke EV, García-Loza GA, Carstens BC, Domínguez CA, Pérez-Ishiwara R, Molina-Freaner F. Geographic isolation and long-distance gene flow influence the genetic structure of the blue fan palm Brahea armata (Arecaceae). Journal of Plant Research 136,277-290.

Larkin I, Meyers EA, Carstens BC, Barrow LN. Predictors of genetic diversity in North American squamates. Heredity 114(2), 131-142.


Provost KL, Yang J, Carstens BC. The impacts of transfer learning, phylogenetic distance, and sample size on big-data bioacoustics. PLoS One 17 (12) e02785222.

Pelletier TA, Parsons D, Decker SK, Crouch S, Franz E, Ohrstrom J, Carstens BC. phylogatR: Phylogeographic data aggregation and repurposing. Molecular Ecology Resources 22(8), 2830-2842.

Fonseca EM, Duckett DJ, Almeida FG, Smith ML, Thomé MCT, Carstens BC. Assessing model adequacy for Bayesian Skyline Plots using posterior predictive simulation. PLoS ONE 17(7): e0269438.

Smith ML, Wallace J, Tank DC, Sullivan J, Carstens BC. The role of multiple Pleistocene refugia in promoting diversification in the Pacific Northwest. Molecular Ecology 31(16), 4402-4416.

Carstens BC, Moshier SP. Giant tree frogs exemplify the promise of integrating multiple types of data in phylogeographic investigations. Molecular Ecology 31(15), 3971-3974.

Ruffley M, Smith ML, Espindola A, Turck D, Mitchel N, Carstens BC, Tank DC, Sullivan J. Genomic evidence of an ancient Inland Temperate Rainforest. Molecular Ecology31(10), 2985-3001.

Parsons D, Duckett DJ, Wieringa J, Pelletier TA, Carstens BC. Analysis of biodiversity data suggest that mammal species are hidden in predictable places. Proceedings of the National Academy of Sciences 119(14), e2103400119.

Carstens BC, Smith ML, Duckett DJ, Fonseca EM, Thomé MCT. Assessing model adequacy leads to more robust phylogeographic inference. Trends in Ecology and Evolution 37(5), 402-410.

Lanna FM, Colli GR, Burbrink FT, Carstens BC. Identifying traits that enable lizard adaptation to different environments. Journal of Biogeography 41(1), 104-116.

Bonatelli IAS, Gehara M, Carstens BC, Colli GR, Moraes EM. Comparative and predictive phylogeography in the dry diagonal of South America: The role of Pleistocene climate change on multi-taxon phylogeography. Molecular Ecology 31(1) 331-342.


Duckett DJ, Sullivan J, Pirro S, Carstens BC. Genomic resources for the North American water vole (Microtus richardsoni) and the montane vole (Microtus montanus). Gigabyte 1, 1-13.

Barrow L, Fonseca EM, Thompson CE, Carstens BC. Identifying predictors of genetic diversity in amphibians with machine learning. Molecular Ecology Resources 21, 2818-2831.

Fonseca EM, Colli G, Werneck F, Carstens BC. Model selection with convolutional neural networks. Molecular Ecology Resources 21, 2818-2831.

Thomé MTC, Carstens BC, Rodrigues MT, Galetti Jr. PM, Haddad CFB. Asynchrony of seasons determines genetic differentiation in a Neotropical toad through neutral and selective mechanisms. Heredity 127, 363-372.

Satler JD, Carstens BC, Garrick RC, Espíndola A. The phylogeographic shortfall in hexapods: A lot of legwork remaining. Insect Systematics & Diversity 5, 1-18.

Morales AE, Fenton MB, Carstens BC, Simmons NB. Myotis evotis and Myotis keenii are two different species. Canadian Journal of Zoology 99, 415-422.

Rankin AM, Anderson FE, Clutts SA, Espíndola A, Carstens BC, Lucid M, Sullivan J. 2021. Comparative phylogeography of two Northern Rocky Mountain endemics: the widespread Anguispira kochi occidentalis and the narrow-range Anguispira nimapuna (Gastropoda: Discidae). Biological Journal of the Linnean Society 133, 817-834.

Wieringa JG, Carstens BC, Gibbs HL. Seasonal distribution and migratory pathways for three species of tree bats: Implications for conservation. PeerJ 9:e11177.

Thomé MTC, Carstens BC, Rodrigues MT, Alexandrino J, Haddad CFB. Rapid diversification patterns in a Neotropical frog supports opposite responses to Pleistocene climates at the Caatinga and Cerrado, regional variation within the Caatinga, and uninterrupted gene flow across a geographic barrier. Journal of Biogeography 48, 1112-1123.

Fonseca EM, Duckett DJ, Carstens BC. P2C2M.GMYC: An R package for identifying model violations under the Generalized Mixed Yule Coalescent model. Methods in Ecology & Evolution 12(3), 487-493.

Thompson CE, Pelletier TA, Carstens BC. Genetic diversity in North American Vertebrates in protected areas. Biological Journal of the Linnean Society 132, 388-398.


Fonseca EM, Colli GR, Werneck FP, Carstens BC. Phylogeographic model selection using convolutional neural networks. bioRxiv 2020.09.11.291856; doi:

Barrow LN, Fonseca EM, Thompson CE, Carstens BC. Predicting amphibian intraspecific diversity with machine learning: Challenges and prospects for integrating traits, geography, and genetic data. bioRxiv 2020.05.03.073049; doi:

Wieringa JG, Nagle J, Nelson DM, Carstens BC, Gibbs HL. 2020. Using trace elements to identify the geographic origin of migratory bats. PeerJ 8:e10082.

Lado P, Smith ML, Carstens BC, Klompen H. 2020. Population genetics structure and demographic history of the lone star tick, Amblyomma americanum (Ixodida: Ixodidae): New evidence supporting old records. Molecular Ecology 29, 2810-2823.

Duckett DJ, Pelletier TA, Carstens BC. Posterior Predictive Simulation for SNAPP Species Tree Inference. PeerJ8:e8271. Supplemental materials.

Smith ML, Carstens BC. Process-based species delimitation leads to the identification of more biologically relevant species. Evolution 74, 216-229.

Weiringa J, Boot M, de Queiroz MD, Duckett DJ, Fonseca EM, Glon H, Hamilton N, Kong S, Lanna FM, Mattingly KR, Parsons D, Smith ML, Stone B, Thompson CE, Zou L, Carstens BC. Does habitat instability structure intraspecific genetic diversity? It’s complicated… Frontiers in Biogeography 12.2, e45377.


Sullivan J, Smith ML, Espíndola AE, Ruffley MR, Rankin AM, Tank DC, Carstens BC. 2019. Integrating life history traits in predictive phylogeography. Molecular Ecology 28, 2062-2073. DOI.

Satler JD, Carstens BC. 2019. The Sarracenia alata pitcher plant system and obligate arthropod inquilines should be considered an evolutionary community. Journal of Biogeography 46, 485-496. DOI.

Rankin AM, Wilke T, Lucid M, Leonard W, Espindola A, Smith ML, Carstens BC, Sullivan J. 2019. Complex interplay of ancient vicariance and recent patterns of geographical speciation in north-western North American temperate rainforests explains the phylogeny of jumping slugs (Hemphillia spp.). Biological Journal of the Linnean Society 127, 876-889. DOI.

Morales AE, Ruedi M, Field K, Carstens BC. 2019. Diversification rates have no effect on the convergent evolution of foraging strategies in the most species-rich genus of bats, Myotis. Evolution 73, 2263-2280.


Pelletier TA, Carstens BC, Tank DC, Sullivan J, Espindola A. Predicting plant conservation priorities on a global scale. Proceeding of the National Academy of Sciences USA. DOI: OSU Press Release. "In This Issue" featuring artwork by Abbie Zimmer

Smith ML, Ruffley MR, Rankin A, Espindola A, Tank DC, Sullivan J, Carstens BC. Testing for the presence of cryptic diversity in the tail-dropper slugs (Genus Prophysaon) using molecular data. Biological Journal of the Linnean Society, 124, 518-532.

Carstens BC, Morales AE, Field KA, Pelletier TA. A global analysis of bats using automated comparative phylogeography uncovers a surprising impact of Pleistocene glaciation. Journal of Biogeography 45, 1795-1805. DOI: 10.1111/jbi.13382.

Morales AE, Carstens BC. Evidence that Myotis lucifugus ‘subspecies’ are five non-sister species, despite gene flow. Systematic Biology, syy010.

Pelletier TA, Carstens BC. Geographic range size and latitude predict population genetic structure in a global survey. Biology Letters14, 20170566.

Ruffley MR, Smith ML, Espíndola AE, Carstens BC, Tank DC, Sullivan J. Model selection reveals complex phylogeography of Red Alder, Alnus rubra, in the Pacific Northwest temperate rainforest. Molecular Ecology, 27, 1012-1024. DOI:10.1111/mec.14491.

Wheeler GL, Carstens BC. Identifying the genomic signal of plant carnivory using gene ontology. Peer J 6:e4322; DOI 10.7717/peerj.4322

Smith ML, Carstens BC. Disentangling the process of speciation using machine learning. bioRxiv.

Carstens BC. Review of Species Concepts in Biology: Historical Development, Theoretical Foundations and Practical Relevance by Frank E. Zachos. Quarterly Review of Biology 93, 36.


Carstens BC, Morales AE, Jackson ND, O'Meara BC. Objective Choice of Phylogeographic Models. Molecular Phylogenetics & Evolution. 116, 136-140. doi:10.1016/j.ympev.2017.08.018 download

Silva-Arias G, Reck-Kortmann M, Carstens BC, Hasenack H, Bonatto S, Freitas L. From inland to the coast: spatial and environmental signatures on the genetic diversity in the colonization of the South Atlantic Coastal Plain. Perspectives in Plant Ecology, Evolution and Systematics 28, 47-57. download

Smith ML, Ruffley MR, Espíndola AE, Tank DC, Sullivan J, Carstens BC. Demographic model selection using random forests and the site frequency spectrum. Molecular Ecology 26, 4562-4573. download

Satler JD, Carstens BC. Do ecological communities disperse across biogeographic barriers as a unit? Molecular Ecology 26, 3533-3545. download

Morales AE, Jackson ND, Dewey TA, O'Meara BC, Carstens BC (2017) Speciation with gene flow in North American Myotis bats. Systematic Biology 66, 440-452. download

Jackson ND, Morales AE, Carstens BC, O'Meara BC (2017) PHRAPL: Phylogeographic Inference using approximate likelihoods. Systematic Biology, in press. doi:10.1093/sysbio/syx001. download

Jackson ND, Morales A, Carstens BC, O'Meara BC (2017) Species delimitation with gene flow. Systematic Biology 66, 799-812. doi:10.1093/sysbio/syw117. download

Gruenstaeudl M, Carstens BC, Santos-Guerra A, Jansen RK (2017) Statistical hybrid detection and the inference of ancestral distribution areas in Tolpis (Asteraceae). Biological Journal of the Linnean Society 121, 133-149. download


Espíndola A, Ruffley M, Smith M, Carstens BC, Tank DC, Sullivan J (2016) Predicting cryptic diversity from phylogeographic, climatic and taxonomic data. Proceedings of the Royal Society B 283, 20161529. download

Pelletier TA, Carstens BC (2016) Comparing range evolution in two western Plethodon salamanders: Glacial refugia, competition, ecological niches, and spatial sorting. Journal of Biogeography, 43, 2237-2249. download

Thomé MCT, Carstens BC (2016) Phylogeographic model selection leads to insight into the evolutionary history of Four-eyed frogs. PNAS 113, 8010-8017. download. A contribution to the Sackler Colloquia In the Light of Evolution X: Comparative Phylogeography.

Satler JD, Carstens BC (2016) Phylogeographic concordance factors quantify phylogeographic congruence among co-distributed species in the Sarracenia alata pitcher plant system. Evolution 70, 1080-1093. download

Carstens BC, Gruenstaeudl M, Reid NM (2016) Community trees: Identifying codiversification in the Páramo dipteran community. Evolution 70, 1105-1119. download

Perez MF, Bonatelli IAS, Moraes E, Carstens BC. Refining phylogeographic inference in two interglacial microrefuge-dwelling cactus species using model-based analysis. Journal of Heredity, 116, 550-557. download

Thomé MCT, Brusquetti F, Sequeira F, Carstens BC, Haddad CFB, Rodrigues MT, Alexandrino J (2016) Recurrent connections between Amazon and Atlantic forests shaped diversity in Caatinga Four-Eyed frogs. Journal of Biogeography43, 1045-1056. download

Sovic MG, Carstens BC, Gibbs HL (2016) Genetic diversity in migratory bats: Results from RADseq data for three tree bat species at a wind farm in northwestern Ohio. PeerJ 4:e1647; doi 10.7717/peerj.1647. download

Gruenstaeudl M, Reid NM, Wheeler G, Carstens BC (2016) Posterior predictive checks of coalescent models: P2C2M, an R package. Molecular Ecology Resources 16, 193–205. download

Satler JD, Zellmer AJ, Carstens BC (2016) Biogeographic barriers drive co-diversification within associated eukaryotes of the Sarracenia alata pitcher plant system. PeerJ 4:e1576; doi 10.7717/peerj.1576. download

Perez MF, Carstens BC, Rodrigues GL, Moraes EM (2016) Anonymous nuclear markers data supporting species tree phylogeny and divergence time estimates in a cactus species complex in South America. Data in Brief 6, 456-460. download

Perez MF, Carstens BC, Rodrigues GL, Moraes EM (2016) Anonymous nuclear markers reveal taxonomical incongruences and long-term disjunction in a cactus species complex with continental-island distribution in South America. Molecular Phylogenetics and Evolution 95, 11-19. doi:10.1016/j.ympev.2015.11.005 download


O'Meara BC, Jackson ND, Morales AE, Carstens BC. (2015) Phylogeographic Inference using approximate likelihoods. bioRxiv.

Bonatelli IAS, Carstens BC, Moraes EM (2015) Using next generation RAD sequencing to isolate multispecies microsatellites for Pilosocereus (Cactaceae) genus. PLoS ONE 10: e0142602. Doi:10.1371/journal.pone.0142602. download

Hird SM, Sanchez C, Carstens BC, Brumfield RT (2015) Comparative gut microbiota of 59 Neotropical bird species. Frontiers in Microbiology 6, 1403. doi: 10.3389/fmicb.2015.01403 download

Pelletier TA, Crisafulli C, Wagner S, Zellmer AJ, Carstens BC. (2015) Historical species distributions predict species limits. Systematic Biology 64, 909-925. download

McVay JD, Flores-Villela O, Carstens BC (2015) Diversification of North American Natricine snakes. Biological Journal of the Linnean Society 116, 1-12. download

Garrick RC, Bonatelli IAS, Morales AE, Pelletier TA, Perez MF, Rice E, Satler JD, Symula RE, Thomé MTC, Carstens BC (2015) The evolution of phylogeographic datasets. Molecular Ecology 24, 1164-1171. download


Espindola A, Carstens BC, Alvarez N. (2014) Comparative phylogeography of mutualists and the effect of the host on the genetic structure of its partners. Biological Journal of the Linnean Society 113, 1021-2035. download

Gavin DG, Fitzpatric MC, Gugger PF, Heath KS, Rodriquez-Sanches F, Dobrowski SZ, Hampe A, Hu FS, Ashcroft MB, Bartlein PJ, Blois JL, Carstens BC, Davis EB, de Lafontaine G, Edwards ME, Fernandez M, Henne PD, Herring RM, Holden ZA, Kong W-S, Liu J, Magri D, Matzke NJ, McGlone MS, Saltre F, Stigall AL, Tsai Y-H E, Williams JW. (2014) Climate refugia: Using fossils, genetics, and spatial modeling to explain the past and project the future of biodiversity. New Phytologist 204, 37-54. download

Pelletier TA, Carstens BC. (2014) Model choice in phylogeography using a large set of models. Molecular Ecology 23, 3028-3043. download

Hird SM, Carstens BC, Cardiff SW, Dittman DL, Brumfield RT. (2014) Sampling locality is more important than taxonomy or ecology in the gut microbiota of the brood-parasitic Brown-Headed Cowbird (Molothrus ater). PeerJ 2; e21. download

Reid NM, Brown JM, Satler JD, Pelletier TA, McVay JD, Hird SM, Carstens BC. (2014) Poor fit to the multi-species coalescent model is widely detectable in empirical data. Systematic Biology 63, 322-333. download


Carstens BC, Satler JD. (2013) The carnivorous plant described as Sarracenia alata contains two cryptic species. Biological Journal of the Linnean Society 109, 737-746. download. Information about primers.

Carstens BC, Brennan RS, Chua V, Duffie CV, Harvey MG, Koch RA, McMahan CD, Nelson BJ, Newman CE, Satler JD, Seeholzer G, Prosbic K, Tank DC, Sullivan J (2013) Model selection as a tool for phylogeographic inference: An example from the willow Salix melanopsis. Molecular Ecology 22, 4014-4028. download

Carstens BC, Pelletier TA, Reid NM, Satler JD (2013) How to fail at species delimitation. Molecular Ecology 22, 4369-4383. download

Salter JD, Carstens BC, Hedin M (2013) Multilocus species delimitation in a complex of morphologically conserved trapdoor spiders (Mygalomorphae, Antrodiaetidae, Aliatypus). Systematic Biology 62, 805-823. download

Carstens BC (2013) Phylogeography. Oxford Bibliographies

Andrew R, Bernatchez L, Bonin Z, Buerkle CA, Carstens BC, Emerson BC, Garant D, Giraud T, Kane NC, Rogers SM, Slate J, Smith H, Sork VL, Stone GN, Vines TH, Waits LS, Widmer A, Rieseberg LH (2013) A Roadmap for Molecular Ecology. Molecular Ecology 22, 2605-2626. download

McVay JD, Carstens BC. (2013) Testing monophyly without well-supported gene trees: Evidence from multi-locus nuclear data conflicts with existing taxonomy in the snake tribe Thamnophiini. Molecular Phylogenetics & Evolution 68, 425-431. download

McVay JD, Carstens BC (2013) Phylogenetic model choice: Justifying a species trees or concatenation analysis. Phylogenetics and Evolutionary Biology 1, 1000114. download

McCormack JE, Hird SM, Zellmer AJ, Carstens BC, Brumfield RT (2013). Applications of next-generation sequencing to phylogeography and phylogenetics. Molecular Phylogenetics and Evolution 66, 526-538. download

Tsai Y-H E, Carstens BC (2013) Assessing model fit in phylogeographic investigations: An example from the North American willow Salix melanopsis. Journal of Biogeography 40, 131-141. download


Reid NM, Carstens BC (2012). Phylogenetic estimation error can decrease the accuracy species delimitation: A Bayesian implementation of the General Mixed Yule Coalescent model. BMC Evolutionary Biology 12, 196, 1-11. download

Carstens BC, Lemmon AR, Moriarty-Lemmon E (2012). The promises and pitfalls of next-generation sequencing data in phylogeography. Systematic Biology 61, 713-715. download

Zellmer AJ, Hanes MM, Hird S, Carstens BC. (2012). Deep phylogeographic structure and environmental differentiation in the carnivorous plant Sarracenia alata. Systematic Biology 61, 763-777. download


Hird SM, Brumfield RT, Carstens BC (2011). PRGMATIC: an efficient pipeline for collating genome-enriched second-generation sequencing data using a ‘provisional reference genome’. Molecular Ecology Resources 11, 743-748. download

Ence DD, Carstens BC (2011). SpedeSTEM: A rapid and accurate method for species delimitation. Molecular Ecology Resources 11, 473-480. download

Koopman MM, Carstens BC (2011) Plant genetic divergence predicts microbial community structure: Molecular phyllogeography of carnivorous pitcher plants. Microbial Ecology 61, 750-758. download


Carstens BC, Dewey TA (2010) Species delimitation using a combined coalescent and information theoretic approach: An example from North American Myotis bats. Systematic Biology 59, 400-414. download

Hird S, Kubatko L, Carstens BC (2010) Replicated subsampling enables accurate species tree estimation in empirical systems. Molecular Phylogenetics & Evolution 57, 888-898. download

Koopman MM, Carstens BC (2010) Conservation genetic inferences in the carnivorous plant Sarracenia alata (Sarraceniaceae). Conservation Genetics 11, 2027-2038. download

Koopman MM, Fuselier DM, Hird S, Carstens BC (2010) Bacterial characterization of the carnivorous Pale Pitcher Plant reveals diverse, distinct, and temporally dependent communities. Applied and Environmental Microbiology 76, 1851-1860. download

Hickerson MJ, Carstens BC, Cavender-Bares J, Crandall KA, Graham CH, Johnson JB, Rissler L, Victoriano PF, Yoder AD (2010) Phylogeography’s past, present, and future: 10 years after Avise 2000. Molecular Phylogenetics & Evolution 54, 291-301. download

Carstens BC, Knowles LL (2010) Navigating the Unknown: Model selection in phylogeography. Molecular Ecology 19, 4581-4582. download

Fleming TH, Murray KL, Carstens BC (2010) Phylogeography and Genetic Structure of Three Evolutionary Lineages of West Indian Phyllostomid Bats. Chapter 5 in Evolution, Ecology, and Conservation of Island Bats, T.H. Fleming and P.A. Racey, eds. Univ. Chicago Press. download


Carstens BC, Stoute HN, Reid NM (2009) An information theoretical approach to phylogeography. Molecular Ecology 18, 4270-4282. download

Koopman MM, Gallagher E, Carstens BC (2009) Isolation and characterization of nine microsatellite loci in the Pale Pitcher Plant Sarracenia alata (Sarraceniaceae). Molecular Ecology Resources 9, 1460-1466. download

Cantrell M, Carstens BC, Wichman HA (2009) X Chromosome Inactivation and XIST Evolution in a Rodent Lacking LINE-1 Activity. PLoS One 4, e6252. download

Kubatko L, Carstens BC, Knowles LL (2009) STEM: Species tree estimation using maximum likelihood for gene trees under coalescence. Journal of Bioinformatics 25, 971-973. download

Eckert AJ, Carstens BC. (2008) Does gene flow destroy phylogenetic signal? The performance of three methods for estimating species phylogenies in the presence of gene flow. Molecular Phylogenetics & Evolution 49, 832-842. download


Carstens BC, Knowles LL (2007) Estimating phylogeny from gene tree probabilities in Melanoplus grasshoppers despite incomplete lineage sorting. Systematic Biology 56, 400-411. download

Carstens BC, Richards CL (2007) Integrating coalescent and ecological niche modeling in comparative phylogeography. Evolution 61, 1439-1454. download

Carstens BC, Knowles LL (2007) Shifting distributions and speciation: genomic resolution of species divergence during rapid climate change. Molecular Ecology 16, 619-627. download

Knowles LL, Carstens BC (2007) Delimiting species without monophyletic gene trees. Systematic Biology 56, 887-895. download

Richards CL, Carstens BC, Knowles LL (2007) Distribution modeling and statistical phylogeography: an integrative framework for generating and testing alternate biogeographical hypotheses. Journal of Biogeography 34, 1833-1845. download

Knowles LL, Carstens BC, Keat ML (2007) Coupling genetic and ecological-niche models to examine how past population distributions contribute to divergence. Current Biology 17. 1-7. download

Brunsfeld SJ, Miller TR, Carstens BC (2007) Insights into the biogeography of the Pacific Northwest of North America: Evidence from the phylogeography of Salix melanopsis (Salicaceae). Systematic Botany 32, 129-139. download

Knowles LL, Carstens BC (2007) Estimating a population-divergence model for statistical phylogeographic tests in montane grasshoppers. Evolution 61, 477-493. download


Carstens BC, Knowles LL (2006) Variable nuclear markers for Melanoplus oregonensis identified from the screening of a genomic library. Molecular Ecology Notes 6, 683-685. download

Carstens BC, Brunsfeld SJ, Demboski JR, Good JD, Sullivan J (2005) Investigating the evolutionary history of the Pacific Northwest mesic forest ecosystem: Hypothesis testing within a comparative phylogeographic framework. Evolution 59, 1639-1652. download

Carstens BC, Degenhardt JD, Stevenson AL, Sullivan J (2005) Accounting for coalescent stochasticity in testing phylogeographic hypotheses: Modeling Pleistocene population structure in the Idaho Giant Salamander Dicamptodon aterrimus. Molecular Ecology 14, 255-265. download

Carstens BC, Bankhead III A, Joyce P, Sullivan J (2005) Testing population genetic structure using parametric bootstrapping. Genetica 124, 71-75. download

Steele CA, Carstens BC, Storfer A, Sullivan J (2005) Testing hypotheses of speciation timing in Dicamptodon copei and Dicamptodon aterrimus (Caudata: Dicamptodontidae). Molecular Phylogenetics & Evolution 36, 90-100. download

Johnson DG, Carstens BC, Sheppard WS, Zack RS (2005) Phylogeny of the leafhopper subgenus Errhomus (Erronus) (Hemiptera: Homoptera: Cicadellidae) based on mitochondrial DNA sequences. Annals of the Entomological Society of America 98, 165-172. download


Carstens BC, Stevenson AL, Degenhardt JD, Sullivan J (2004) Testing nested phylogenetic and phylogeographic hypotheses in the Plethodon vandykei species group. Systematic Biology 53, 781-792. download

Carstens BC, Sullivan J, Davalos LM, Larsen PA, Pedersen SC (2004) Exploring population genetic structure in three species of Lesser Antillean bats. Molecular Ecology 13, 2557-2566. download

Carstens BC, Lundrigan BL, Myers P (2002) A phylogeny of the Neotropical nectar-feeding bats (Chiroptera: Phyllostomidae) based on morphological and molecular data. Journal of Mammalian Evolution 9, 23-53. download


We make inferences about the history of species and ecological communities using genetic, environmental and morphological data. We conduct investigations in a range of organisms: bats, insects, carnivorous plants, salamanders, willows, spiders, snails, slugs and viruses.

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